The MKIII runs it close, but I'd have to say the 300 f2.8. Without that I could never have hoped to produce a high enough standard of work to move on from where I was.
I found (when they worked) my Canon kit of MKIII with 400MM and MKIII with 135M was pretty good.
However i have noticed a dramatic increase in my keeper rate since the switch to Nikon (D3S 400MM,70+200VRII), mainly with the AF system acquiring (sp?) and locking on to a target better. Though having talked to guys shooting with the MKIV they too have seen a big increase in the AF performance
My 3rd replacement Canon MKIII and my 400mm F/2.8 L IS was stunning until Lehmanns screwed the lens up but then the move to Nikon made me think that the Canon was in fact just a heavy point and shoot kit.
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